Eamonn Holmes and Ruth Langsford have been replaced on their This Morning slot – and replaced by another popular duo.
The husband and wife duo, have been presenting the Friday shows for 14 years.
They have been doing the gig since 2006, and are said to be “furious and upset” over the move.
But in a shake-up of presenters, the couple will instead be given the job of hosting the main programme for six weeks in the summer holidays, when main hosts Phillip Schofield and Holly Willoughby take a break.
The Friday shows will now go into the hands of Alison Hammond and Dermot O’Leary.
Mum-of-one Alison, 45, has been part of the This Morning “family” for years as a roving reporter and is a firm favourite with fans.
She and former X Factor host O’Leary, 47, have previously been paired as stand-in presenters – most recently last month – and bosses have been hugely impressed by their chemistry, reports Mirror Online.
“They have real warmth,” an insider said. “It’s time for a change and they are perfect.
“Eamonn is pretty furious and everyone feels for him and Ruth, they are well liked on the team, but sometimes tough decisions have to be made.”
Eamonn, 60, has confided in recent weeks that he feared being too “pale, stale and male” for the award-winning ITV show, which has drawn criticism for not having enough diversity among the presenters.
Earlier this summer ITV chief executive Carolyn McCall announced a Diversity Acceleration Plan after the channel was criticised for having an entire day with only one non-white face.
Holmes also left bosses fuming earlier this year when he was rapped by Ofcom for “ill-judged” comments regarding the conspiracy theories surrounding 5G masts spreading coronavirus.
The watchdog said his remarks, in which he said that mainstream media should not automatically slap down such claims, “risked undermining viewers’ trust in advice from public authorities and scientific evidence”.
Ruth, also 60, is expected to remain on Loose Women, where internal research has shown she remains hugely popular with viewers.
The changes, brought about by show boss Martin Frizell following feedback from the regular audience, are expected to start in the New Year.
Hammond, who found fame on Big Brother and is known for her fun approach, was given the job of guest editing the entire show for one day last month.
Prior to her takeover – which she co-hosted with O’Leary – she told fans there would be only jolly items during her day in charge.
“It will just be a joy!” she said. “There will be nothing about lockdown or coronavirus.
“There’ll be a lot of recipes and food… and maybe a little bit of fashion and fun. It’s going to be great and I’m excited to be the boss.”
Dermot and Alison are expected to finalise their new contracts in the coming weeks.